Nathan Fillion as Malcolm "Mal" Reynolds
A former sergeant on the losing side of the Unification War, he struggles to survive free and independent of the Alliance.
Gina Torres as Zoe Washburne
A former corporal who fought under Mal in the war, and Wash's wife. She is fiercely loyal to Mal, whom she addresses as "sir".
Summer Glau as River Tam
River is a
seventeen-year old psychic genius. She and her brother are taken in by
the crew of Serenity after he rescues her from an Alliance Academy where
she was subjected to medical experimentation and brainwashing. The
Alliance's pursuit of River acts as the film's motive. However, more
abstractly the film is the "story of Mal as told by River".
Morena Baccarin as Inara Serra
A
Companion who formerly rented one of Serenity's shuttles. In one of the
Operative's traps, Mal is reunited with Inara at her training house, and
the two escape back to Serenity.
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